В работе использована классификация, разработанная Глезер с соавт. (1988), дополненная новыми родами (Cox, 1987, 1988 Round et al., 1990 Lange-Bertalot, Metzeltin, 1996 Lange-Bertalot, 1997, 1999 Schiller


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492
ISSN 0868
-
8540
;L3�
�;38O. 2010. ". 20. ! 4
Alg
ologia. 2010. V. 20. N 4
5
8
2.26+5
8
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BACILLARIOPHYTA
.
1. .
NAVICULACEAE
K

TZ
.

.
⠀. ⤀

11
3


,
.

.
6
3



.









Nupela
Vyverman
et
Comp

re

Prestauroneis
Bruder
et
Medlin
.
7
ϒ.



Й

pH
.


:


,
. ,
Bacillariophyta
,
,

, ϒ.







ә
ϙ
Д

Gutwi

sky
, 1
8
97;
Ko

odziejczyk
,
1916⤀
ߐ
ә



,
1999
;
., 2004

.







:
,

.







Й


.

. ␓


.


.
297 , Ӟ 5160
2


Г ,








.


.













⠀᧐, , 197
8
;
, 㞎,
1979;

., 2006⤀
.

20
,

Ӟ
,
,

, 1939;
߷
,
1968; ., 1998⤀. ᑓә
, .. , ..
-
.!. C;8:�2A:89, !..
5=:0;, ".. 8E5520, 2010
�2K5 40==K5 � D�;@5
Bacillariophyta
5;0@CA8
493
⠀1999⤀


887
,
.

.
309
.





.




,

ϒ

.





,




.

2006
-
200
8
. 7
,
⠀ 2
-
6
⤀ :






2 3⤀,


-
⠀ 4 5⤀
0,5



Ӟ ⠀
6⤀
.

1


Ӟ


Й
䞧,
7

10
, Й



.


⠀, 1975⤀
.


ӳ ⠀
Nikon
Eclipse
E
600⤀ ⠀
JSM
-
25
S
⤀.


ϒ, 㔙
.
⠀19
88

,




Cox
, 19
8
7
, 19
88
;
Round
et
al
.,
1990;
Lange
-
Bertalot
,
Metzeltin
, 1996;
Lange
-
Bertalot
, 1997, 1999;
Schiller
,
Lange
-
Bertalot
, 1997;
Moser
et
al
., 199
8

.






116

,

.

.
6
3


.

,

,
.
Adlafia
Lange
-
Bertalot
A
.
bryophila
⠀Petersen⤀ Lange
-
Bertalot



.
I,
9


=
Navicula bryophila
Petesen,
N.
maillardii
Germain⤀
.
13,
3
.,
3
В
.
A
.
suchlandtii
⠀Hust
.
⤀ Lange
-
Bertalot



.
I,
10

⠀=
Navicula suchlandtii
Hust
.

.


16
.,
4,
5
В.,
27
А 10 .
Amphipleura
Ktz
.
A
.
ruti
lans
⠀Trentep
.
⤀ Cleve



.
II,
16


=
Conferva rutilans
Trentep
.
,
Berkeleya rutilans
⠀Trentep
.
⤀ Grun
ow

.


19,
3
.,
3,
3
В
.
.!. C;8
:�2A:89, !.. 5=:0;, ".. 8E5520
494
Caloneis
Cleve
C
.
biconstrictoides
Levkov



.
II,
1
8

.

36
-
44,
7
.,
10
-
12
В.,
19
А 10
.
C
.
hyalina
Hust
.



.
II,
17

.

17,
3
-
1
8
.,
5
-
5,
3
В.,
21
-
24
А 10 .
C
.
silicula
var.
elliptica
Freng
.



.
II,
19

.

22,
7
-
36
.,
7,
7
-
10
В.,
21
-
23
А 10 .
C
.
silicula
var.
minuta
⠀Grun
ow
⤀ Cl
eve



.
II,
13

.


30


.,
6

В
.,
23
А

10

.
Chamaepinnularia
Lange
-
Bertalot et
Krammer
Ch
.
krookii
⠀Grun
ow
⤀ Lange
-
Bertalot et
Krammer



.
I,
14


=
Navicula
krookii
Gr
un
ow
,
N.
ignobilis
Krasske,
Pinnularia ignobilis
⠀Krasske
⤀ Cleve
-
Euler,
P.
krookii
⠀Grun
ow
⤀ Cleve⤀
.

24
.,
6
В.,
20
А 10 .
Craticula
Grun
ow
C
.
a
mbigua

Eh
r
enb.
⤀ D.G. Mann



.
III,
3


=
Navicula ambigua
Ehr
enb.
,
N.
cuspidata
var.
ambigua

Ehr
enb.
⤀ Cle
ve⤀
.

49,3
-
61,
3

.,
14
-
16,
7
В.,
17
-
1
8
А 10 .
Diadesmis
Ktz
.
D
.
confervaceae
Ktz
.



.
II
I
,
14


=
Navicula confervaceae
⠀Ktz
.

Grun
ow
,
N.
confervaceae
var.
hungarica
Grun
ow
,
N. confervaceae
var.
peregrina
Grun
ow
,
Diadesmis peregrina
W. Sm.

.

12
-
20
.,
5,3
-
7,
3
В.,
20
-
21
А 10 .
Diploneis
Ehr
enb.
D
.
petersenii
Hust
.



.
II
I
,
3
1

.


14,
7
.,
6
В.,
24
А 10 .
Eolimna
Lange
-
Bertalot et
Schill
.
E
.
subminuscula
⠀Mang
.
⤀ Moser, Lange
-
Bertalot et
Metzel
tin



.
I,
8


=
Navicula subminuscula
Mang
.
,
N.
luzonensis
Hust
.
,
N.
demissa
Hust
.
,
N.
frugalis
Hust
.
,
N.
vaucheriae
Petersen sensu Hust
.
,
N.
pervarva
Hust
.
sensu Cholnoky

.


12
.,
5
В.,
1
8
А 10 .
Fallacia
A.J. Stickle et
D.G. Mann
F
.
insociabilis
⠀Krasske⤀ D.G. Mann



.
I
II,
2
8


=
Navicula insociabilis
Krasske,
N.
fritschii
Lund,
N.
natalensis
Cholnoky
⤀.


10
.,
3,
7
В.,
24
А 10 .
F
.
pygmaea

K

tz
.

D
.
G
.
Mann


.
III,
3
2


=
Navicula pygm
aea
Ktz
.,
N.
minutula
W. Sm.
,
N.
rotundata
Hantzsch fide Grun
ow
in Van Heurck 1
88
0,
N.
hudsonis
Grun
ow
,
Diploneis hudsonis
Cleve
⤀.

19,3
-
31,
3
.,
8
-
12
В.,
24
-
30
А 10 .
F
.
subhamulata
⠀Grun
ow
⤀ D.G. Mann



.
III,
2
7


=
Navic
ula
subhamulata
Grun
ow

.

10,
7
-
1
8
.,
4,7
-
5,
7
В.
�2K5 40==K5 � D�;@5
Bacillariophyta
5;0@CA8
495
Frustulia
Rabenh
.
F
.
krammeri
Lange
-
Bertalot et
Metzeltin



.
IV,
4

.


102,
7
.,
20,
7
В.,
24
А 10 .
Geissleria
Lange
-
Bertalot et
Metzeltin
G
.
decussis
⠀Hust
.
⤀ Lange
-
Bertalot et
Metzeltin



.
I,
13


=
Navicula
decussis

str
.
,
N.
terebrata
Hust
.
,
N.
exiguiformis
Hust
.
,
N.
exiguiformis
f.
capitata
Hust
.

.

1
8
,
7
-
26
.,
6,
3
-
8
В.,
16
-
1
8
А 10 .
Gyrosigma
Hass
.
G
.
exile
⠀Grun
ow
⤀ Reimer



.
IV,
3

.


100
.,
13,
3
В.,
17
А 10 .
G
.
obtusatum
⠀Sull
.
et
Wormley⤀ Boyer



.
IV,
2

.

8
2
-
126,
7
.,
13,3
-
1
8
,
7
В.,
1
8
-
19
А 10 .
G
.
spencerii
⠀Quekett⤀ Griff
.
et
Henfrey



.
IV,
1


=
Navicula spencerii
Quekett,
Pleurosigma ktzingii
Grun
ow
,
P.
spencerii
var.
ktzingii
⠀Grun
ow
⤀ Grun
ow
,
Gyrosigma ktzingii
⠀Grun
ow
⤀ Cleve⤀
.

88
-
113,
3
.,
13
,
3
-
14
В.,
20
-
21
А 10 .
Hippodonta
Lange
-
Bertalot, Metzeltin et
Witkowsk
i
H
.
arkonensis
Lange
-
Bertalot, Metzeltin et
Witkowski



.
III,
3
0

.


14
.,
4
В.,
14
А 10 .
Kobayasiella
Lange
-
Bertalot
K
.
parasubtilissima
⠀Kobayasi et
Nagumo⤀ Lange
-
Bertalot



.
I,
4


=
Navicula parasubtilissima
Kobay
asi et
Nagumo,
Kobayasia parasubtilissima
⠀Kobayasi
et
Nagumo⤀ Lange
-
Bertalot⤀
.

29,3
-
30,
7
.,
4,
7
В.
Luticola
D.G. Mann
L
.
goeppertiana
⠀Bleisch⤀ D.G. Mann



.
II,
1
5


=
Navicula mutica
var.
goeppertiana
M. Bleisch⤀
.

1
8
,
7
-
34
.,
7,3
-
9,
3
В.,
1
8
-
19
А 10 .
Mayamaea
Lange
-
Bertalot
M
.
atomus
var.
permites
⠀Hust
.
⤀ Lange
-
Bertalot



.
III,
2
6


=
Navicula
permites
Hust
.
,
N.
peratomus
Hust
.
,
N.
atomus
var.
permites
⠀Hust
.
⤀ Lange
-
Bertalot,
N.
atomus
var.
alcimon
ica
Reich
.
,
Mayamaea atomus
var.
alcimonica
⠀Reich
.
⤀ Reich
.
,
M.
permites
⠀Hust
.
⤀ Bruder et
Medlin
⤀.


9,
3
.,
3,
3
В.
M
.
fossalis
⠀Krasske⤀ Lange
-
Bertalot



.
III,
29


=
Navicula fossalis
Krasske⤀
.

9,
3
-
10
.,
3,
3
-
4
В.
,
14
А 10 .
Navicula
Bory
N
.
antonii
Lange
-
Bertalot



.
III,
15


=
N
.
antonii
var.
grunowii
Lange
-
Bertalot⤀
.

11,3
-
20,
7
.,
5,3
-
6,
7
В.,
14
-
15
А 10 .
.!. C;8
:�2A:89, !.. 5=:0;, ".. 8E5520
496
N
avicula
associata
Lange
-
Bertalot



.
III,
13


=
N
.
reichar
dtiana
var.
crassa
Lange
-
Bertalot et
Hof
f
.

.

1
8
-
20
.,
6
-
7,
3
В.,
17
-
1
8
А 10 .
N
.
cryptofallax
Lange
-
Bertalot et
Hof
f
.



.
III,
7

.


22
.,
5,
3
В.,
17
А 10 .
N
.
cryptotenelloides
Lange
-
Bertalot



.
III,
16, 17

.

10,7
-
16,
7
.,
3,
7
-
4
В.,
19
А 10 .
N
.
exilis
Ktz
.



.
III,
14


=
N
.
cryptocephala
var.
exilis
Grun
ow

.

19,3
-
22,
7
.,
5,
3
-
6
,
7
В.,
16
-
1
8
А 10 .
N
.
hofmanniae
Lange
-
Bertalo
t



.
III,
12

.

30,
7
-
32
.,
7,
3
-
8
В.,
17
-
1
8
А 10 .
N
.
libonensis
Schoeman



.
III,
20


=
N
.
viridula
var.
pamirensis
Hust
.
,
N.
schubartii
var.
africana
P.A.
Arch
ibald

.

32
-
33,
3
.,
6
-
6,
7
В.,
13
-
14

10 .
N
.
moscalii
Metzeltin, Witkowski et
Lange
-
Bertalot



.
III,
1
8


=
Frustulia oblonga
Ktz
.

.

17,3
-
24,
7
.,
6,3
-
7,
3
В.,
14
-
16
А 10 .
N
.
oligotraphenta
Lange
-
Bertalot et
Hof
f
.


.
III
,
19

.


27,
3
.,
7,
3
В.,
16
А 10 .
N
.
rhynchotella
Lange
-
Bertalot



.
III,
4

.

4
8
,6
-
50,
7
.,
12
-
12,
7
В.,
11
-
12
А 10 .
N
.
slesvicensis
Grun
ow



.
III,
10


=
N
.
viridula
var.
slesvicensis
⠀Grun
ow

Van Heurck⤀,
N
.
trivialis
Lange
-
Bertalot
.

30,7
-
53,
3
.,
10
-
10,
7
В.,
9
-
10
А 10 .
N
.
stankovicii
Hust
.



.
III,
8
, 9
⤀.

32,
7
-
36,
7
.,
6
В.,
15
А 10 .
N
.
upsaliensis
⠀Grun
ow
⤀ Perag
.



.
III,
5, 6


=
N
.
meni
sculus
var.
upsaliensis
Grun
ow
in Van Heurck 1
88
0,
N.
menisculus
var.
upsaliensis
Grun
ow
in
Cleve et
Grun
ow
1
88
0,
N
.

gastrum
var.?⤀
upsaliensis
in Cleve et
Mll
.
1
88
1⤀
.

21,3
-
45,
3
.,
9,
3
-
10,
7
В.,
13
-
14
А 10 .
N
.
wiesneri
Lan
ge
-
Bertalot



.
III,
21


=
N
.
heufleri
f.
minuta
Grun
ow

.

15,3
-
23,
3
.,
3,7
-
5,
3
В.,
13
А 10 .
Nupela
Vyverman et
Compre
N
.
wellneri
⠀Lange
-
Bertalot⤀ Lange
-
Bertalot



.
I,
5
-
7


=
Navicula
wellneri
Lange
-
Bertalot⤀
.


13,6
-
14,
5
.,
4,5
-
7,
3
В.,
23
-
24
А 10 .
Parlibellus
Cox
P
.
protractoides
⠀Hust
.
⤀ Witkowski, Lange
-
Bertalot et
Metzeltin



.
I,
15


=
Navicula protractoides
Hust
.

.


23,
3
.,
6,
7
В.,
19
А 10 .
Pinnul
aria
Ehr
enb.
P
.
anglica
Krammer



.
II,
10

.


42
.,
8
В.,
12
А 10 .
�2K5 40==K5 � D�;@5
Bacillariophyta
5;0@CA8
497
P
innularia
grunowii
Krammer



.
II,
5

.

37,
3
-
42
.,
8
-
8
,
7
В.,
11
-
12
А 10 .
P
.
isselana
Krammer



.
II,
8


=
P
.
subrup
estris
var.
parva
Krammer,
P.
rupestris
var.
cuneata
Krammer⤀
.


26
,
7
.,
7,
3
В.,
11
А 10 .
P
.
neohalophila
Kulikovskiy, Genkal,
Mikheyeva
nov. stat., nov. nom.



.
II,
2
-
4
⤀.
Basi
onym:
P
.
rhombarea
var.
halophila
Krammer 2000
, Diatoms of Europe,
Vol. 1. p. 217, fig. 54: 6
-
11.

32,7
-
5
8
,
7
.,
10
-
14
В.,
10
А 10 .

P
.
rhombarea
Krammer

߷
,


K
.


Krammer
,
2000⤀
,
,
ߐ
,
, ϙ
, А
10 .
P
innularia
rhombarea
var
.
halophila

-
В , Ϟ, В
А 10 ⠀10
-
1
1, 9
-
10⤀.
ϒ В

, .
P
innularia
rhombaundulata
Kulikovskiy, Genkal nov. stat., nov. nom.
Basi
onym:
P
.
rhombarea
var.
undulata
Krammer 2000, Diatom
s of Europe,
Vol. 1. p. 217, fig. 56: 1
-
3.
ᐔ , В
-
ϙ, В ϙ , В Ϟ.
P
.
rhombavariarea
Kulikovskiy, Genkal nov. stat., nov. nom.
Basi
onym:
P
.
rh
ombarea
var.
variarea
Krammer 2000, Diatoms of Europe,
Vol. 1. p. 21
8
, fig. 56: 4
-
6.

, Ϟ, В .
P
.
brevicapitata
⠀Krammer⤀ Kulikovskiy, Genkal
nov. stat.
Basi
onym:
P
.
rhombarea
var.
brevicapitata
Krammer 2000, Diatoms of
Europe, Vol. 1. p. 21
8
, fig. 56: 7
-
11.
ᐔ В ,
В Ϟ.
P
.
biundulata
⠀O. Mll.

Kulikovskiy, Genkal nov. stat.
Replaced synonym:
P
.
microstauron
var.
biundulata
O. Mll
.
1
8
9
8
, For
-
schungsber. Biol. Stat. Pln. Vol. 6, p. 25.
Synonym:
P
.
microstauron
f.
biundulata
⠀O. Mll
.
⤀ Hust
.
1930, p. 320,
fig. 5
8
3;
P
.
rhombarea
var.
biundulata
⠀O
. Mll
.
⤀ Krammer 2000, p. 76, fig. 56: 12
.
,
»ô , ķ Ŵ ,
Ư Ƿų.
P
.
neomajor
Krammer


İ
.
II,
1

.
ɯ

165,
3
.,
26,
7
.,
8
ʷ 10 .
.!. C;8
:�2A:89, !.. 5=:0;, ".. 8E5520
498
P
innula
ria
obscura
Krasske


İ
.
II,
12

.
ɯ

15,
3
.,
4
.,
9
ʷ 10 .
P
.
latarea
Krammer


İ
.
II,
6

.
ɯDZ
46,7
-
49,
3
.,
8
,
7
.,
10
-
11
ʷ 10 .
̻ ͷ °
P
.
latarea
var
.
thermophila
Krammer
,
Ƿº

,
ķ »ô
Ŵ

.
αô
ǵ Ƿıű.
P
.
thermophila
⠀Krammer⤀ Kulikovskiy, Genkal nov. stat.
Basi
onym:
P
.
latarea
var.
thermophila
Krammer
2000, Diatoms of Eur
ope,
Vol. 1. p. 2
24
, fig.
8
0
: 1
-
6
.
P
.
parvulissima
Krammer


İ
.
II,
9

.
ɯDZ
42,7
-
45,
3
.,
8
,7
-
9,
3
.,
9
ʷ 10 .
P
innularia
perspicua
Krammer


İ
.
II,
7

.
ɯDZ
45,
3
-
66
.,
10,7
-
11,
3
.,
11
-
12
ʷ 10 .
P
.
s
inistra
Krammer


İ
.
II,
11

.
ɯ

2
8
.,
4
.,
12
ʷ 10 .
Placoneis
Mereschkowsky
P
.
elginensis
⠀Greg
.
⤀ Cox


İ
.
I,
3


=
Pinnularia elginensis
Greg
.

.
ɯ

27,
3
.,
8
,
7
.,
11
ʷ 10 .
P
.
ignorata
⠀Schim
anski⤀ Lange
-
Bertalot


İ
.
I,
12


=
Navicula ignorata
Schimanski,
N.
dicephala
f.
cuneata
M. Mller ex Foged 1977,
N.
elginensis
var.
cuneata
⠀M. Mller ex Foged⤀ Lange
-
Bertalot⤀
.
ɯDZ
20,
7
-
26
.,
8
.,
12
ʷ 10 .
P
.
paraelginens
is
Lange
-
Bertalot


İ
.
I,
11

⠀=
⠀?⤀
Placoneis dicephala

Ehrenb.
⤀ Mereschkowsky⤀
.
ɯDZ
22,
7
-
27,
3
.,
7,
3
-
8
.,
12
-
13
ʷ 10 .
P
.
subgastriformis
⠀Hust
.
⤀ Cox


İ
.
I,
2


=
Navicula subgastriformis
⠀Hust
.

Cox⤀
.
ɯ

29,
3

.,
10,
7
.,
14
ʷ 10 .
Prestauroneis
Bruder et
Medlin
P
.
integra
⠀W. Sm.
⤀ Bruder


İ
.
I,
1


=
Pinnularia integra
W. Sm.
,
Navicula integra
⠀W. Sm.
⤀ Ralfs⤀
.
ɯDZ
34,
7
.,
9,
3
.,
1
8
ʷ 10 .
Sellaphora
Mereschkowsky
S
.
cf.
insolita
⠀Manguin ex Kociolek et Reviers⤀ Hamilton et
Antoniades


İ
.
III,
22


=
Navicula insolita
Manguin ex Kociolek et
Reviers⤀
.
ɯDZ
22
-
34
.,
9,
3
-
10
.,
19
-
21
ʷ 10 .
S
.
lanceolata
D.G. Mann et
S. Droop


İ
.
III,
2
3

.
ɯDZ
25,
3
-
2
8
.,
6,7
-
7,
3
.
S
.
obesa
D.G. Mann et
M.M. Bayer


İ
.
III,
24

.
ɯDZ
29,3
-
32,
7
.,
8
,
7
-
10
.,
22
ʷ
10 .
�2K5 40==K5 � D�;@5
Bacillariophyta
5;0@CA8
499
Stauroneis
Ehr
enb.
S
.
amphicephala
Ktz
.


İ
.
III,
2

.
ɯDZ
54
-
54,
7
.,
12
.,
19
ʷ 10 .
S
.
borrichii
⠀Petersen⤀ Lund


İ
.
I
II,
25


=
Navicula borrichii
Petersen,
Stauroneis thienemannii
f.
simplex
Hust
.

.
ɯ

1
8
.,
4,
7
.,
19
ʷ 10 .
S
.
gracilis
Ehr
enb.


İ
.
III,
1

.
ɯ

70
.,
14,
7
.,
17
ʷ 10 .
S
.
reichardtii
Lange
-
B
ertalot, Cavacini, Tagliaventi et
Alfinito


İ
.
III,
11

.
ɯdz
37,
3
.,
9,
3
.,
21
ʷ 10
.
߰
Ƿ
ô ´ ǵDZ
Ƿıű, ´
Ƶ

.
௷ Ƶ ƴ ᴴ ƼǷ ǷŴű


Mann
, 1999
⤀. 䳽
. 峽 ⠀
Mann
et
al
.,
2004


. ߻ ⠀
Droop
et
al
.
,
2000

Ƿ ư
dz

ʷ Ƿ ǷŴű , Ƿ
ǵ
氱 ¯ 紾
´ , ǻ
ᱴ þ
ķ.
η
请 ʷ ƴ ķ , Ƿ
ǵDZ Ƶ ͷ
,
Ƿ
´
óϯ
ͷ . 黱 请
ʷ ưű ʷ ¶

Ư
ı


ǵDZ Ƿ
⠀괽dz,
1993
⤀.

ô


´ ǵDZ Ƿıű , Ƿ´ °
Ľô

Ǵ.
黱 Ʋ
. ɹ
-



Ͱ 촰
Ƶ


Nupela

Prestauroneis
.
鴻º

Ʋ
鳸Ƴ η 깱 ⠀
Vyverman
,
Comp

re
, 1993
⤀.


Nupela
ǰϲ Ƿ 4
0

,
Ƿ


dz




,







Lange
-
Bertalot
,
Moser
, 199
4;
Lange
-
Bertalot
, 1999;
Kusber
,
Metzeltin
, 200
1
;
Monnier
et
al
.,
2003;
Potapova
et
al
., 2003
;
Siver
at
al
.,
2007
⤀.
뱾 请
dz ⠀
Van
de
Vijver
et
al
.
, 2002;
Monnier
et
al
., 2003
⤀.
Nupela
wellneri
DZ dz,
滶 泽DZ
ᴵ; ɴ Ǵ Ľô ŴǷ ⲳ
ķ Ǵ Ƿ 谴ǯ, ó
-
泽DZ ᴵ ⠀
Lange
-
Bertalot
,
Krammer
, 19
8
7;
Potapova
et
al
., 2003⤀.


Pre
stauroneis


Bruder
,
Medlin
200
8

ͷ汲Ƿ
洽Ƿ



.
γ

P
.
integra

º

,
Ƿº


dz Ƿ ⠀
Krammer
,
Lange
-
Bertalot
,
19
8
6
⤀.
.!. C;8
:�2A:89, !.. 5=:0;, ".. 8E5520
5
00
ʷ
dz ý

Hippodonta
arkoensis
.

°
Ƿ
쳰DZ


dz 괻

Lange
-
Bertalot
et
al
.,
1996;
Lange
-
Bertalot
, 2001

.

,







,
Ŵ ͷ
,
Ǽ
ıǷ
-
Ŵ


. Ľô
dz 请 ƴ
⽴Ƿ ͷ.
Stauroneis
reichardtii
,
Pinnularia
latarea

P
.
perspicua


´ . 鴻º
° ɳ


ǯDZ ䷵, 껴 ʱ
Svalbard

Lange
-
Bertalot
et
al
., 2003;
Van
de
Vijver
et
al
., 2004

.
߹
ǷDZ ƾ 쳹

Krammer
,
2000

.
Navicula
parasubtilissima
°
Ʋ


N
.
subtilissima
P
.
T
.
Cleve

Kobayasi
,
Nagumo
, 19
88

.
黱 请
° dz, 踱
N
.
subtilissima
ñ
,
²


DZ dz
⠀.
,
Krammer
,
Lange
-
Bertalot
, 19
8
6
:
Fig
. 79: 22
-
27
⤀.

ķ

-

N
.
subtilissima
ǷǷ

Krammer
,
Lange
-
Bertalot
,
19
8
6⤀
.
鷼ô 诱 °


Kobaya
si
a
Lange
-
Bertalo
t

Lange
-
Bertalot
, 199
6

,

Kobay
a
siella

Lange
-
Bertalot
,
1999⤀
.
߰
촰Ƶ

N
.
subtilissima
dz¹
..
ʴ
-


1999⤀, Ƿ ý ñ,


ǵ

K
.
parasubtilissima
, ķ
º ʳ
-
º 请 汷
.
ʷ Ƴű,
,


Ʋ
ʷ

Ƶ
Navicula
atomus
K

tz
.

=
Mayamaea
atomus

K

tz
.

Lange
-
Bertalot

⠀ʴ, 1999⤀.

,
请 ° 淺
ķ


º ǵ

M
.
atomus
var
.
permites
. 黱 请, Ƿ
ƴ Ƿ Ǵ ⠀Ĵ ., 1951⤀

N
.
atomus
ƴ 请 ,
M
.
atomus
var
.
permites

Kulikovskiy
, 2006

.


Sellaphora
pupula
-
com
plex


ͷ 请
dz ⠀
Mann
,
2001,
2004⤀
. ƶ ǷŴű
,

dz¹


Ƶ
ǵ

Sellaphora
lanceolata

S
.
obesa
.
黱º

Sellaphora
cf
.
insoli
ta
, ý
,
dz
S
.
bacillum
, ô
Ľô
º
. ɹ.

.
̻ .
-


请 ǵ dz
S
.
bacillum

Krammer
,
Lange
-
Bertalot
, 19
8
6⤀.

°
ô

Jahn
et
al
.
,
200
8

.
̻
Ͱ 촰
Ƶ


50
,

.
ɹ
:
Aneumastus
rostratus

Hust
.

Lange
-
Bertalot

Navicula tuscula
var.
rostrata
Hust
.
,
N.
tusculoides
Cleve
-
Euler⤀,
A
.
stroesei

str
.
⤀ Ma
nn et
Stickle

Navicula t
uscula
var.
stroesei

str
.
,
�2K5 40==K5 � D�;@5
Bacillariophyta
5;0@CA8
501
N.
pseudotuscula
Hust
.

,
A
.
sphaerophora
⠀Ehr
enb.
⤀ Pfitzer

Navicula
sphaerophora
Ehr
enb.

,
Caloneis amphisbaena
⠀Bory⤀ Cleve

Navicula
amphisbaena
Bory⤀,
C
.
lancettula
⠀Schulz⤀ Lange
-
Bertalot et
Witkowski

C.
aemula
var.
lance
ttula
Schulz,
C.
bacillum
var.
lancettula
⠀Schulz⤀ Hust
.
⤀,
C
.
limosa
⠀Ktz
.
⤀ Patrick

Navicula limosa
Ktz
.
,
N.
gibberula
Ktz
.
,
N.
trochus
Ehr
enb.
,
N.
schumanniana
Grun
ow
,
N.
schumanniana
var.
biconstricta
⠀Grun
ow
⤀ Reich
.
,
Caloneis schumanniana
⠀Grun
ow

Cleve

,
C
.
tenuis
⠀Greg
.

Krammer

Pinnularia tenuis
Greg
.
,
P.
gracillima
Greg
.
⤀,
Craticula cuspidata
⠀K
tz
.
⤀ D.G. Mann

Frustulia cuspidata
Ktz
.
,
Navicula cuspidata
⠀Ktz
.

Ktz
.
,
N.
cuspidata
var.
heribaudii
M. Perag
.

,
Diploneis alpina
Meist
.

D.
dombl
ittensis
var.
subconstricta
A.
Cleve,
D.
didyma
var.
bavarica
f.
typical
Mayer,
D.
subconstricta
⠀A. Cleve⤀ Cleve
-
Euler⤀
,
D
.
elliptica
⠀Ktz
.
⤀ Cleve

Navicula elliptica
K
tz
.

,
D
.
oculata
⠀Brb
.
⤀ Cleve

Navicula oculata
Brb
.

,
D
.
parma
Cleve
,
D.
subovalis
Cleve
,
Eolimna minima
⠀Grun
ow
⤀ Lange
-
Bertalot

Navicula minima
Grun
ow
,
N.
minutissima
Grun
ow
,
N.
atomoides
Grun
ow
,
N.
minima
var.
atomoides
⠀Grun
ow
⤀ Cleve,
N.
tantula
Hust
.

,
F
rustulia
saxonica
Rabenh
.

F.
rhomboids
auct. partim.,
F.
rhomboids
var.
saxoni
ca
⠀Rabenh
.
⤀ Pfitz
.

,
F
.
vulgaris
⠀Thw
.
⤀ De Toni

Schizonema vulgare
Thw
.
,
Frustulia vulgaris
var.
elliptica
Hust
.

,
Geissleria paludosa
⠀Hust
.
⤀ Lange
-
Bertalot morphotype 2

Navicula lagerstedtii
var.
palustris
Hust
.
,
N.
ignota
var.
palustris
⠀Hust
.
⤀ Lund,
N.
paludosa
Hust
.
⤀,
G
.
schoenfeldii
⠀Hust
.
⤀ Lange
-
Bertalot et
Metzeltin

Navicula schoenfeldii
Hust
.
⤀,
G
.
similis
⠀Krasske⤀
Lange
-
Bertalot et
Metzeltin

Navicula similis
Krasske⤀
,
Gyrosigma
attenuatum
⠀K
tz
.
⤀ Rabenh
.

Frustulia attenuata
Ktz
.

,
Hippodont
a capitata
⠀Ehr
enb.
⤀ Lange
-
Bertalot, Metzeltin et
Witkowski

Navicula capitata
Ehr
enb.
⤀,
H
.
lueneburgensis
⠀Grun
ow
⤀ Lange
-
Bertalot, Metzeltin et
Witkowski

Navicula hungarica
var.
lueneburgensis
Grun
ow
,
N.
capitata
var.
lueneburgensis
⠀Grun
ow
⤀ Patrick⤀
,
Lu
ticola ventriconfusa
Lange
-
Bertalot

Navicula mutica
var.
ventricosa
⠀Ktz
.
⤀ Grun
ow
sensu Hust
.
1966,
N.
neoventricosa
Hust
.
1966 ⠀excl. Basionym⤀,
N.
mutica
var.
ventricosa
⠀Ktz
.

Grun
ow
sensu Krammer et
Lange
-
Bertalot 19
8
6

,
Navicula cari
Ehr
enb.

N.
ci
ncta
var.
cari
⠀Ehr
enb.
⤀ Cleve⤀,
N
.
capitatoradiata
Germ
.

N
.
cryptocephala
var.
intermedia
Grun
ow
,
N.
sali
narum
var.
intermedia
⠀Grun
ow
⤀ Cleve⤀,
N
.
cincta
⠀Ehr
enb.
⤀ Ralfs

Pinnularia cincta
Ehr
enb.
,
N
.
heufleri
Grun
ow
,
N.
inutilis
Krasske,
N.
tumida
Bock
⤀,
N
.
cryptocephala
Ktz
.

N
.
cryptocefalsa
Lange
-
Bertalot⤀,
N
.
cryptotenella
Lange
-
Bertalot

N.
tenella
Brb
. ex K
tz
.
,
N.
radiosa
var.
tenella
⠀Brb
.
ex K

tz
.
⤀ Van Heurck⤀,
N
.
oblonga
⠀Ktz
.
⤀ Ktz
.
,
N.
reichardtiana
Lange
-
Bertalot

Navicula exilis
K
tz
.
sensu Lange
-
Bertalot
1979⤀,
N
.
reinhardtii
⠀Grun
ow
⤀ Grun
ow

Stauroneis reinhardtii
Grun
ow
⤀,
N.
tripunctata
⠀O.F. Mll
.
⤀ Bory

Vibrio tripunctata
O.F. Mll
.
,
Navicula gracilis
Ehr
enb.
,
Schizonema neglectum
Thw
.

,
N
.
veneta
Ktz
.

N.
cryptocephala
var.
venet
a
⠀Ktz
.
⤀ Rabenh
.
,
N.
cryptocephala
var.
subsalina
Hust
.

,
N
.
viridula
⠀Ktz
.
⤀ Ehr
enb.

Frustulia viridula
Ktz
.
,
Pinnularia silesiaca
Bleisch ex
.!. C;8
:�2A:89, !.. 5=:0;, ".. 8E5520
502
Fresenius 1
8
62⤀,
Neidium ampliatum
⠀Ehr
enb.
⤀ Krammer

Navicula ampliata
Ehr
enb.
,
Neidium iridis
var.
ampliata
⠀Ehr
enb.
⤀ Cleve⤀,
N
.
binodis
⠀Ehr
enb.

Hust
.

Navicula binodis
Ehr
enb.
⤀,
N
.
bisulcatum
⠀Lag
.
⤀ Cleve

Navicula
bisulcata
Lag
.
⤀,
N
.
dubium
⠀Ehr
enb.
⤀ Cleve

Navicula dubia
Ehr
enb.
⤀,
Parlibellus protracta
⠀Grun
ow
⤀ Witkowski, Lange
-
Bertalot et
Metzeltin

Navicu
la protracta
Grun
ow
⤀,
Pinnularia borealis
Ehr
enb.
,
P.
globiceps
Greg
.
,
P
.
microstauron
⠀Ehr
enb.
⤀ Cleve

Stauroptera microstauron
Ehr
enb.
,
S.
parva
Ehr
enb.
,
Navicula bicapitata
var.
hybrida
Grun
ow

,
P
.
viridis
⠀Nitzsch⤀
Ehr
enb.

Bacillaria viridis
Nitzsch⤀,
Placoneis clementis
⠀Grun
ow
⤀ Cox

Navicula clementis
Grun
ow
⤀,
P
.
gastrum
⠀Ehr
enb.
⤀ Meresckowsky

Pinnularia gastrum
Ehr
enb.
⤀,
P
.
placentula
⠀Ehr
enb.
⤀ Heinzerling
*

Navicula
placentula
⠀Ehr
enb.
⤀ Ktz
.
,
Pinnularia placentula
Ehr
enb.
⤀,
P
.
pseudanglica
⠀Lange
-
Bertalot⤀ Cox

Navicula pseudanglica
Lange
-
Bertalot⤀,
Sellaphora
bacillum
⠀Ehr
enb.
⤀ D.G. Mann

Navicula bacillum
Ehr
enb.
⤀,
S
.
pupula
⠀Ktz
.

Mereschkowsky

Navicula pupula
Ktz
.
⤀,
Stauroneis
cf.
kriegeri
Patrick

S.
anceps
var.
capitata
Perag
.
,
S.
pygmae
a
Krieg
.

.
Ƿ汲DZ ¶´
ķ
ϯ ͽ´, dzͱ´ Ƶ
.

洷滳ͱ
Ƿ ʳǯǴ 请 Ƿ ´
dz ķ

Krammer
,
L
ange
-
Bertalot
, 19
8
6;
Lange
-
Bertalot
, 2001

.
௱ ʳǯDZ ó
Ƶ
. ɹ
-

, ô

蹯ͱ
,

Ƿ
Ƿ

, Ƿ

䷵, ô 巽淰

Kara
yeva
,
Gen
kal
, 1993
;
Lange
-
Bertalot
, 199
8
;
Werum
,
Lange
-
Bertalot
, 2004
;

Ƿ
, 2005
;
ưǷDZ, 2008;
ưǷDZ, ߴDZ, 200
8
⤀.

,
űͱ´ Ƶ
,

Chamaepinnularia
krooki
,
Frustulia
krammeri
,
F
.
saxonica
,
Kobayasiel
la
parasubtilissima
,
Pinnularia
sinistra
.
鴻º
ǵ ı
淯ͽ

Lange
-
Bertalot
,
Genkal
, 1999

.

Ƿ ϯ
Ƿ
űͱ
Ͱ
ͳ潷 ķ, Ƿ


ϯ ķ Ƿ
⠀ưǷDZ, 2007

.
F
rustulia
krammeri


汷 ,
ɱı ɴ
DZ
,
ķ Ƿ
ű

Ƿ 谴ǯ
-
*
ʷº ͷ汱 诱
N
avicula
gastrum

Ehr
.⤀
K

tz
.
var
.
latiuscula
Grun
.
ķ ǰϹ Ư Ƿų DZ,
ʳǯdz. αô ǵ Ƿıű:
Placoneis latiuscula
⠀Grun
.
⤀ Kulikovskiy, Genkal nov. com
b., nov. stat.
Basionym:
Navicula gastrum
⠀Ehr
.
⤀ Ktz
.
var.
latiuscula
Grun
.
in Cleve et Grun
.
, K. Sv. Vet.
Akad. Handl., 1880: 31.
Synonym:
Placoneis placentula
f.
latiuscula
⠀Grun
.
⤀ Bukhtiyarova 1995, Algologia, Vol. 5⠀4⤀,
p. 420.
�2K5 40==K5 � D�;@5
Bacillariophyta
5;0@CA8
503
, ô ͳ潷
ķ

Lange
-
Bertalot
,
2001

.


dz º
Ƿ Ͱ . ɹ.
γ űͱ Ƿ
Ͱ Ƿ


ᴰ Ƶ

Ƿ
-

,



ķ ͳ潷 ķ

ij DZ
ķ , ķ

dz
ô
.
㋉儍
ǯ ᴵ . ɹ ¶
µǷ
ķ泯
ư


116

,

.

.
11
3
Ͱ DZ
,
6
3


촰Ƶ
.
Ľô´
ʳǯ ͽ, , 澳
dz
űͱº Ƿ Ͱ 淲 ǰϲ
,
ʳǯ´


ͳ潷 ķ.


ķ ¸ ͱ Ǵ ䷵Ƿ ͷ
ƽ

⠀滳 0
8
-
04
-
90007⤀ 촰ƵǷ
İdz

Ƿ ͷ ƽ
⠀滳
0
8

-
104⤀.
M
.
S
.
Kulikovskiy
1
,
S
.
I
.
Genkal
1
,
T
.
M
.
Mik
heeva
2
1
I.D. Papanin Institute of Biology of Inland Waters, Russian Academy of Sciences,
Settle of Borok, Nekouzsky District, 152742 Yaroslavl Region, Russia
2
Belarusian State University,
4, Nezavisimosti Av., 220030 Minsk, Belarus Republic
NEW DATA
FO
R
THE
BACILLARIOPHYTA
FLORA
OF
BELARUS.
1. FAMILY
NAVICULACEAE
KTZ.
In the phytoplankton of the Svisloch River ⠀Belarus Republic⤀ 113 diatom taxa of species and
infraspecific rank new for this river have been revealed. Among them 63 taxa are firs
t cited for Belarus flora.
Genera
Nupela
Vyverman
et
Compre et
Prestauroneis
Bruder
et
Medlin were not
recorded in Belarus before.
In the paper 7 new taxonomical combinations are suggested. All revealed taxa are known to prefer mesotrophic
and eutrophic a
lkaline water bodies.
Keywords:
Belarus, Svisloch River,
Bacillariophyta
, flora, new taxa, taxonomic combinations.
ȗɖ
..
Ž Ƚɽ ˽ ˽Ͻ ˽½
// Ľ̽ϽȽ
Ž
.

.: , 1975.

.
8
7
-
8
9.
.!. C;8
:�2A:89, !.. 5=:0;, ".. 8E5520
504
μЖ ..
ڽ ʽϽ ½ Ž ˽

Bacillariophyta

// ´ɽ.

1995.

5
,
4.

. 417
-
424.
ӑԗ ..
ľ Ͻ Ž ˽
Centr
ophyceae
: .
.
...


., 1993.

41 .
ӎ֑ .., З .., ˗ԗЖ ..
.
Ž ˽ //
Ž ˽ ⠀½ ʽ⤀. .
II
, . 1.

ݼ.
:

, 19
88
.

. 31
-
35.
חڑɗ ..
, .., ЖԼɗ
-
їБԖ .., ޑԖ ..
Ž
˽
//
̾ Ͻ ˽ . . 4.

.
:
ݽ.

, 1951.

61
8
.
ЏԖ ..
Ƚ ˽ Ž Ž.

.
.
Ͼ.

.
.

, 196
8
.

25 .
̎ԖԼ ..
Ž ˽ ½ ÿϽ ֽɽ ˽½
: . . Ͼ. . .

ݼ., 2007.

24 .
̎ԖԼ ..
Ž ˽ ½ Ͻ: ˽
˽ ɽ
̿ĽϽ //
½ :
ĽϽ, ½ɽ, .

.

.
˽. ½
-
½.

˽
,
200
8
.

.
179
-
1
8
1.
̎ԖԼ .., Լ ..
Ž ⠀
Bacillariophyta

½
̾
. Ͽ ⠀ҽϿ⤀ // ½
.
.

200
8
.

2
.

. 101
-
111.
˼͑ ..
´ɽ
. ʽϽ ɽ.

:
,
1999.

396 .
˼͑ .., ӗɐԖ ..
ϾϽ Ľ ˽

Ͻ ÿ // ľ

Ľ. .

1979
.

15
,
1.

. 53
-
63.
˼͑ .., Η .., ԗ ..
.
½
-
Ͻ


Ͻ ½.



: ɿ, 199
8
.

196 .
Η
.., ˼͑
..
˽ ˽˽ ĽϽ
н ÿ ˽ʽ Ž
// . ɽ.
-
.

197
8.

2
,
2
.

.
33
-
3
8
.
Η .
.
, ˼͑ .., ԗ .., Ԗ ..
½ ľ

Ľɽ

н
.
ɽ ⠀

̾⤀ //

.
6
-
.
. ½.

1
8
-
19 2006
.,


.





:
,
2006.
Жɖ .
. ÿ
Ž Ž.

.
:
,
1939.

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Ͻ
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´ɽ.

2004.

14
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4.

. 413
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17
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505
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19
88
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3
, N
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1993.

23
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88
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Warszawa:
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2000.

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703 p.
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Stuttgart
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8
6.

8
76
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.
Kulikovskiy M.S.
To study of morphology and distribution of certain species from genera
Mayamaea
Lange
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Fistulifera
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⤀ from Russia
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2006.

8
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P. 325
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r W.
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H., Metzeltin D.
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2001.

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5
8
5
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.
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1997.

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8
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199
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2
.

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axa aus
Achnanthes
Bory
⠀sensu lato⤀
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.
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.

1999.

6
.

P. 270
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2
8
0.
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Bertalot H., Cavacini P., Tagliaventi N., Alfinito S.
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species in rock pools and other ephemeral waters //
Ibid
.

2003.

12
.

43
8
p.
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-
Bertalot
H., Genkal S.I.
Diatoms from Siberia
I //
Ibid
.

1999
.

12
.

P. 1
-
265.
Lange
-
Bertalot H., Krammer K.
Bacillariaceae, Epithemiaceae, Surirellaceae
. Ne
ue und wenig
bekannte taxa, neue kombinationen und s
ynonyme sowie Bemerkungen und
Ergnzungen
zu den
Nav
iculaceae
//
Bibl
.
Diatom
.

19
8
7.

15
.

P.
1
-
29.
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-
Bertalot H., Metzeltin D.
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.
Diatom
.

1996.

2
.

P. 1
-
390
.
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:�2A:89, !.. 5=:0;, ".. 8E5520
506
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-
Bertalot H., Metzeltin D., Witkowski A.
Hippodonta
gen. nov.

Umschreibung und
Begrndung
einer neuen Gattung der
Naviculaceae
//
Ibid
.

1996.

4
.

S
. 247
-
275.
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Bertalot H., Moser G.
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. Monographie der Gattung
//
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.
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1994.

29
.
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Bertalot H.
Kobayasiella
nov. nom. ein neuer Gattungsname fr
Kobayasia
Lange
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Ber
talot
1996 // Ibid.

1999.

6
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Bertalot H.
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sensu stricto. 10 genera separated from
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sensu lato,
Frustulia
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2001.

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526 p.
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1999.

3
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2001.

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Mann D.G., McDonald S.M., Bayer M.M
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The
Sellaphora pupula
species complex

Bacillariophyceae
⤀: mo
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and mating data provide
evidence for five new species // Phycologia.

2004.

43
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iogeography //
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Antarctic regi
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2002.

46
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20
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.
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.

2004.

13
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11.02.09
.. ½õĸ

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